Try this. Pull both of the plugs on the pc board keeping the tank plugged in. Now test the continuity of the grounds from one cable plug to the other.
It sounds to me like the head is oscillating when the reverb is turned up. The 3-15 cabinet is lower in impedance than the two, so the power amp may be effected by the lower output load.
On the back panel where the speaker jacks are, are the blue wires that connect the speaker jacks to the pc board pushed close to the tank?